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Scientific publications
Anderson, M.D. & Richardson, P.R.K. (2005). The physical and thermal characteristics of aardwolf dens.
South African Journal of Wildlife Research, 35(2): 147-153.
Funston, P.J., Mills, M.G.L., Richardson, P.R.K. & van Jaarsveld, A.S. (2003). Reduced dispersal
and opportunistic territory acquisition in male lions (Panthera leo). Journal of Zoology, London.
259: 131-142.
de Villiers, M.S., Richardson, P.R.K. & van Jaarsveld, A.S. (2003). Patterns of coalition
formation and spatial association in a social carnivore, the African wild dog Lycaon pictus.
Journal of Zoology, London. 260: 377-389.
Molteno, A.J, Sliwa, A. & Richardson, P.R.K. (1998). The role of scent marking in a free-ranging,
female black-footed cat (Felis nigripes).
Sliwa, A., Richardson, P.R.K. 1998. Responses of aardwolves, Proteles cristatus, Sparrman 1783,
to translocated scent marks. Animal Behaviour. 56:137-146.
Williams, J.B., Anderson, M.D. & Richardson, P.R.K. 1997. Seasonal differences in field metabolism,
water requirements, and foraging behaviour of free-living aardwolves in South Africa. Ecology 78:
2588-2602.
De Villiers, M.S., van Jaarsveld, A.S., Meltzer, D.G.A. & Richardson, P.R.K. (1997). Social dynamics
and the cortisol response to immobilisation stress of the African wild dog, Lycaon pictus. Hormones
and Behavior. 31: 3-14.
Anderson, M.D., Williams, J.B. & Richardson, P.R.K. 1997. Laboratory metabolism and evaporative water
loss of the aardwolf. Physiological Zoology 70: 464-469.
Van Jaarsveld, A.S., Richardson, P.R.K. & Anderson, M.D. 1995. Post-natal growth and sustained
lactational effort in the aardwolf: life history implications. Functional Ecology. 9: 492-497.
De Villiers, M.S., Meltzer, D.G.A., van Heerden, J., Mills, M.G.L., Richardson, P.R.K. & van
Jaarsveld, A.S. (1995). Handling-induced stress and mortalities in African wild dogs (Lycaon pictus).
Proceedings of the Royal Society of London, B 262: 215-220.
Richardson, P.R.K. & Anderson, M.D. (1993). Physical capture of the aardwolf Proteles cristatus. In:
The capture and care manual: capture, care, accommodation and transportation of wild African mammals. Editor: A.A. McKenzie. Wildlife Decision Support Services, and South African Veterinary Foundation, South Africa.
Richardson, P.R.K. & Anderson, M.D. (1993). Chemical capture of the aardwolf Proteles cristatus. In:
The capture and care manual: capture, care, accommodation and transportation of wild African mammals. Editor: A.A. McKenzie. Wildlife Decision Support Services, and South African Veterinary Foundation, South Africa.
Jackson, T.P., Skinner, J.D. & Richardson, P.R.K. (1993). Some costs of maintaining a perennial territory in the springbok, Antidorcas marsupialis. African Journal of Ecology. 13: 242-254.
Richardson, P.R.K. (1993). The function of scent marking in territories: a resurrection of the
intimidation hypothesis. Transactions of the Royal Society of South Africa. 48: 195-206.
Anderson, M.D. & Richardson, P.R.K. (1992). Remote immobilization of the aardwolf. South African
Journal of Wildlife Research. 22: 26-28.
Anderson, M.D., Richardson, P.R.K. & Woodall, P.F. (1992). Functional analysis of the feeding
apparatus and digestive anatomy of the aardwolf (Proteles cristatus). Journal of Zoology, London.
228: 423-434.
Richardson, P.R.K. (1991). Territorial significance of scent marking during the non-mating season in
the aardwolf Proteles cristatus (Carnivora: Protelidae). Ethology. 87: 9-27.
Richardson, P.R.K. (1990). Scent marking and territoriality in the aardwolf (Proteles cristatus). In
"Chemical Signals in Vertebrates" p. 378-387. Editors: Macdonald, D.W., Muller-Schwarze, D.,
Natynczuk, S.E. Oxford: Oxford University Press.
Koehler, K.E. & Richardson, P.R.K. (1990). (Proteles cristatus). Mammalian Species No. 363: 1-6.
Apps, P., Viljoen, H.W., Richardson, P.R.K. & Pretorius, V. (1989). Volatile components of the anal
gland secretion of the aardwolf (Proteles cristatus). Journal of Chemical Ecology. 15: 1685-1688.
Richardson, P.R.K. & Coetzee, M. (1988). Mate desertion in response to female promiscuity in the
socially monogamous aardwolf. South African Journal of Zoology. 23: 306-308.
Richardson, P.R.K. (1987). Aardwolf mating system: Overt cuckoldry in an apparently monogamous mammal.
South African Journal of Science. 83: 405-410.
Richardson, P.R.K. (1987). Food consumption and seasonal variation in the diet of the aardwolf
(Proteles cristatus) in Southern Africa. Zeitschrift fur Saugetierkunde. 52: 307-325.
Richardson, P.R.K. (1987). Aardwolf: the most specialized myrmecophagous mammal? South African
Journal of Science. 83: 643-646.
Richardson P.R.K., Mundy, P.J., Plug, I. (1986). Bone crushing carnivores and the significance of
osteodystrophy in griffon vulture chicks. Journal of Zoology, London. 210: 23-43.
Richardson, P.R.K. (1985). The social behaviour and ecology of the aardwolf, Proteles cristatus
(Sparrman, 1783) in relation to its food resources. D.Phil. thesis, University of Oxford.
Richardson, P.R.K. (1984). The scavenging behaviour of vultures in southern Africa. Proceedings of
the Fifth Pan-African Ornithological Congress: 331-353.
Richardson P.R.K. (1983). An improved darting system for immobilizing smaller mammals in the wild.
South African Journal of Wildlife Research. 13: 51-54.
Liversidge, R., Richardson, P.R.K. & Gubb, A. (1981). The sooty falcon Falco concolor in the southern
Kalahari. Bulletin of the British Ornithological Society. 101: 268-270.
Richardson, P.R.K. (1980). Carnivore damage to antelope bones and its archaeological implications.
Palaeontologia Africana. 23: 109-125.
Richardson, P.R.K. (1980). The natural removal of ungulate carcasses, and the adaptive features of
the scavengers involved. M.Sc. thesis, University of Pretoria, Pretoria.
Popular Articles
Richardson, P.R.K. (2002). The aardwolf. In: "The Encyclopaedia of Mammals: 1" p. Editor: Macdonald,
D.W., George Allen & Unwin, London.
Richardson, P.R.K. (1997). Aardwolf Proteles cristatus. In "The complete book of Southern African
Mammals" p. 168. Editors: Mills, M.G.L. & Hes, L. Struik Winchester, Cape Town.
Richardson, P.R.K. (1992). Pangolin: A perambulating pine cone. Signature Magazine.
Richardson, P.R.K. (1990). Termitenjager Eerdwolf: Seine waffe ist die Zunge. Das Tier. 10:14-17.
Richardson, P.R.K. (1990). The lick of the aardwolf. Natural History. 4: 78-85.
Richardson, P.R.K. (1988). Aardwolf, silent and solitary watcher of the night. African Wildlife. 42:
271-275.
Richardson, P.R.K. (1987). Protele. Une Hyene Paradoxale. L'Univers du Vivant. 23: 24-33.
Richardson, P.R.K. (1986). Aardwolf: African werewolf of the savannas. Quagga 14: 6-10.
Richardson, P.R.K. & Bearder, S.K. (1984). The aardwolf. In: "The Encyclopaedia of Mammals: 1"
p.158-159. Editor: Macdonald, D.W. George Allen & Unwin, London.
Supervision of Theses
Funston, P. J. (1999). Predator-prey relationships between lions and large ungulates in the Kruger
National Park. Ph.D. thesis, University of Pretoria, Pretoria.
Swart, J. (1998). Foraging behaviour of the Cape pangolin Manis temminkii in the Sabi Sand Wildtuin.
M.Sc. thesis, University of Pretoria, Pretoria.
Garai, M.E. (1997). The development of social behaviour in translocated juvenile African elephants
Loxodonta africana (Blumenbach). Ph.D. thesis, University of Pretoria, Pretoria.
Sliwa, A. (1996). A functional analysis of scent marking in the aardwolf, Proteles cristatus
(Sparrman, 1783). Ph.D. thesis, University of Pretoria, Pretoria.
de Villiers, M.S. (1995). Adrenocortical stress responsiveness and social dynamics in captive wild
dogs (Lycaon pictus). Ph.D. thesis, University of Pretoria, Pretoria.
Jackson, T.P. (1995). The role of territoriality in the mating system of the springbok Antidorcas
marsupialis (Zimmerman 1780). Ph.D. thesis, University of Pretoria, Pretoria.
Anderson, M.D. 1994. The influence of seasonality and quality of diet on the metabolism of the
aardwolf, Proteles cristatus (Sparrman 1783). M.Sc. thesis, University of Pretoria, Pretoria.
Howard, P. (1992). The ethology, sociology and interspecific interactions of the yellow mongoose,
Cynictis penicillata, in the Karoo. M.Sc. thesis, University of Pretoria, Pretoria.
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